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Field notes · chapter 47

The Respiratory System

pp. 1044–1062 · Raven Part VII

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Partial pressures you should memorize (mm Hg)

6 rows
LocationP_O₂P_CO₂
Dry air at sea level1590.2
Alveolar gas10540
Blood entering the lungs (from body)4046
Blood leaving the lungs (to body)10040
Blood after unloading at tissues4046
Air at 6000 m altitude~80

Lung volumes

7 rows
MeasureValue / definition
Tidal volume~500 mL per resting breath
Anatomical dead space~150 mL — conducting airways, no exchange
Inspiratory reserve volumeExtra air you can inhale beyond tidal
Expiratory reserve volumeExtra air you can force out beyond tidal
Vital capacityTidal + IRV + ERV = 4.6 L men, 3.1 L women
Residual volumeNever exhaled; prevents collapse
Total lung capacityVital capacity + residual volume

How each group breathes

6 rows
GroupOrganMechanismEfficiency note
Bony fishGills under an operculumBuccal + opercular pumping, or ram ventilationCountercurrent (180°) — most efficient of all
ArthropodsTracheae and tracheolesAir enters valved spiraclesAir reaches cells directly, no blood transport
AmphibiansSimple saclike lungs + skinPositive pressure — pushes air inLow surface area; skin supplements
ReptilesLungsNegative pressure via rib cageDry skin blocks cutaneous exchange
MammalsAlveolar lungsNegative pressure via diaphragm + intercostalsHuge area (80 m²) but two-way flow mixes stale air
BirdsParabronchi + air sacsOne-way flow over two cyclesCrosscurrent (90°) — best of the land vertebrates

How carbon dioxide is carried

3 rows
FormShareDetail
Bicarbonate (HCO₃⁻)72%Made in red blood cells by carbonic anhydrase; the main plasma buffer
Bound to hemoglobin20%Carbaminohemoglobin — binds protein, not iron, so no competition with O₂
Dissolved in plasma8%Simple physical solution

Hemoglobin saturation

3 rows
ConditionP_O₂SaturationUnloaded
Arterial blood100 mm Hg97%
Venous blood at rest40 mm Hg75%22%
Venous blood during exercise20 mm Hg35%62%

Exam traps

10 pairs

Questions get built out of near-misses. If you can state each difference in one sentence, you will not lose those points.

Oxygen is limiting in FISHvsCarbon dioxide is limiting in MAMMALS

Water oxygen swings with depth, temperature, and flow, so fish monitor O₂. Air oxygen is essentially constant, so mammals monitor CO₂ instead. Expect this comparison on the exam.

Countercurrent (fish, 180°)vsCrosscurrent (birds, 90°)

Countercurrent is the most efficient arrangement that exists. Crosscurrent is second — better than a mammal lung, not as good as a gill.

Positive pressure breathingvsNegative pressure breathing

Amphibians PUSH air in with the mouth floor. Reptiles, birds, and mammals PULL air in by expanding the chest. Only amphibians push.

Light on percentage vs pressure at altitudevsThe oxygen fraction never changes

Air is 20.95% oxygen at sea level AND on Everest. What falls is total pressure, and therefore P_O₂. Do not say "there is less oxygen in the air."

Bohr effect shifts the curve RIGHTvsA left shift means tighter binding

Lower pH and higher temperature = shift RIGHT = hemoglobin releases oxygen more easily. Exercising muscle creates both conditions on purpose.

HemoglobinvsMyoglobin

Hemoglobin: 4 chains, 4 O₂, in blood. Myoglobin: 1 chain, 1 O₂, in muscle, and it has HIGHER affinity — which is why it holds oxygen back as a last reserve.

CO₂ binds hemoglobin proteinvsO₂ binds the heme iron

Different sites, so they do not compete. But CO₂ binding changes hemoglobin's shape and lowers its oxygen affinity anyway.

HyperventilationvsHeavy breathing during exercise

Hyperventilation means P_CO₂ drops abnormally low. Exercise breathing matches a real metabolic demand and keeps blood gases normal — that is not hyperventilation.

Vital capacityvsTotal lung capacity

Vital capacity is everything you can move. Total lung capacity adds the residual volume you can never exhale.

Trachea (vertebrate)vsTracheae (arthropod)

Same word, unrelated structures — not homologous. One is a windpipe, the others are air tubes reaching cells directly.