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Field notes · chapter 42

The Nervous System

pp. 924–952 · Raven Part VII

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Key terms

120 terms
Sensory (afferent) neuron§42.1
Carries a signal from a receptor to the CNS
Motor (efferent) neuron§42.1
Carries a signal from the CNS out to a muscle or gland
Interneuron (association neuron)§42.1
Neuron entirely within the CNS that links other neurons
Central nervous system (CNS)§42.1
Brain plus spinal cord — the command center
Peripheral nervous system (PNS)§42.1
All sensory and motor neurons outside the brain and spinal cord
Somatic nervous system§42.1
Voluntary motor control of skeletal muscle
Autonomic nervous system§42.1
Involuntary control of smooth muscle, cardiac muscle, and glands
Sympathetic division§42.1
Autonomic division for fight or flight
Parasympathetic division§42.1
Autonomic division for rest and repose
Cell body§42.1
Enlarged neuron region holding the nucleus; integrates dendrite input
Dendrite§42.1
Branched extension that receives input and carries it toward the cell body
Dendritic spine§42.1
Small projection adding receiving surface area to a dendrite
Axon§42.1
The single output fiber carrying impulses away from the cell body
Glial cells (neuroglia)§42.1
Support cells; ~1/10 the size of neurons but ~10× more numerous
Schwann cell§42.1
PNS glial cell that wraps one axon segment in myelin
Oligodendrocyte§42.1
CNS glial cell that myelinates segments of several axons
Myelin sheath§42.1
Insulating wrap of compacted glial membrane around an axon
Node of Ranvier§42.1
Gap in the myelin sheath, every 1–2 µm, where action potentials fire
White matter§42.1
CNS regions of myelinated axons
Gray matter§42.1
CNS regions of unmyelinated cell bodies and dendrites
Nerve§42.1
PNS bundle of myelinated axons, like wires in a cable
Astrocyte§42.1
Star-shaped glial cell; helps build the blood–brain barrier, clears excess K⁺
Blood–brain barrier§42.1
Capillary barrier restricting what passes from blood into the CNS
Ependymal cell§42.1
Ciliated glial cell lining brain cavities; makes cerebrospinal fluid
Microglia§42.1
Immune glial cells acting as CNS macrophages
Resting membrane potential§42.2
Steady voltage of an unstimulated neuron; -40 to -90 mV, standard -70 mV
Na⁺/K⁺ pump§42.2
Active transporter moving 3 Na⁺ out for every 2 K⁺ in
Leakage channel§42.2
Ion channel that is always open; sets the resting potential
Gated channel§42.2
Ion channel that opens only in response to a stimulus
Ligand-gated channel§42.2
Channel opened by a chemical signal; produces graded potentials
Voltage-gated channel§42.2
Channel opened by a change in membrane potential; produces action potentials
Equilibrium potential§42.2
Voltage at which diffusional and electrical forces on one ion balance
Nernst equation§42.2
E_K = 58 mV · log([K⁺]out/[K⁺]in); computes one ion’s equilibrium potential
Graded potential§42.2
Small, stimulus-proportional, local change in potential that fades with distance
Depolarization§42.2
Membrane potential becomes less negative (e.g. -70 → -65 mV)
Hyperpolarization§42.2
Membrane potential becomes more negative (e.g. -70 → -75 mV)
Repolarization§42.2
Return toward the resting potential after depolarization, driven by K⁺ efflux
Summation§42.2
Combining of graded potentials to amplify or cancel each other
Threshold§42.2
Minimum depolarization (~-55 mV) that opens voltage-gated Na⁺ channels
Action potential§42.2
All-or-none spike from -70 mV to about +35 to +40 mV, lasting ~1 ms
Undershoot§42.2
Brief hyperpolarization after repolarization because K⁺ channels close slowly
Absolute refractory period§42.2
Interval when Na⁺ channels are inactivated and the cell cannot fire at all
Relative refractory period§42.2
Interval when the cell can fire, but only a smaller action potential
Saltatory conduction§42.2
Impulse jumping node to node along a myelinated axon; much faster
Giant axon§42.2
Very large-diameter invertebrate axon (squid) giving speed without myelin
Synapse§42.3
Junction where a neuron communicates with another cell
Presynaptic / postsynaptic cell§42.3
Sender and receiver across a synapse
Electrical synapse§42.3
Gap-junction connection passing potential changes directly and fast
Chemical synapse§42.3
Junction using a chemical messenger across a cleft; most vertebrate synapses
Synaptic cleft§42.3
Narrow gap between pre- and postsynaptic cells, visible only by electron microscope
Synaptic vesicle§42.3
Axon terminal sac storing neurotransmitter for exocytosis
Neurotransmitter§42.3
Chemical released by the presynaptic cell to affect the postsynaptic cell
Neuromuscular junction§42.3
Chemical synapse between a motor neuron and a muscle fiber
Acetylcholine (ACh)§42.3
Transmitter at the neuromuscular junction and many neuron-to-neuron synapses
Acetylcholinesterase (AChE)§42.3
Extremely fast enzyme that clears ACh by cleaving it in the cleft
Glutamate§42.3
Major excitatory neurotransmitter of the vertebrate CNS
GABA / glycine§42.3
Inhibitory transmitters that open Cl⁻ channels and hyperpolarize the membrane
Catecholamine§42.3
Tyrosine-derived biogenic amine: epinephrine, norepinephrine, dopamine
Dopamine§42.3
Movement-related transmitter; loss causes Parkinson, excess linked to schizophrenia
Serotonin§42.3
Tryptophan-derived amine regulating sleep and mood; target of SSRIs
Neuromodulator§42.3
Neuropeptide with subtler, longer-term postsynaptic effects
Substance P§42.3
Neuropeptide released in the CNS by pain-sensing sensory neurons
Endogenous opiates§42.3
Enkephalins and endorphins, which block pain via the opiate receptors
Nitric oxide (NO)§42.3
Gaseous transmitter made from arginine on demand; cannot be stored in vesicles
EPSP§42.3
Excitatory postsynaptic potential — a depolarization toward threshold
IPSP§42.3
Inhibitory postsynaptic potential — a hyperpolarization away from threshold
Synaptic integration§42.3
Summing EPSPs and IPSPs at the cell body to decide whether to fire
Spatial summation§42.3
EPSPs from many different synapses adding together at one time
Temporal summation§42.3
Repeated EPSPs from one synapse adding within about 15 ms
Habituation§42.3
Loss of responsiveness after prolonged constant stimulation
Tolerance§42.3
Need for more drug as the postsynaptic cell down-regulates its receptors
Nerve net§42.4
Diffuse web of neurons in cnidarians with no central control
Tract§42.4
Bundle of axons forming an information highway inside the CNS
Hindbrain (rhombencephalon)§42.4
Cerebellum, pons, and medulla oblongata; coordinates motor reflexes
Midbrain (mesencephalon)§42.4
Contains the optic tectum; reflexes involving eyes and ears
Forebrain (prosencephalon)§42.4
Diencephalon plus telencephalon; dominant in later vertebrates
Cerebellum§42.4
Hindbrain region coordinating movement, limb position, and balance
Medulla oblongata§42.4
Hindbrain region for autonomic functions and sensory nuclei
Diencephalon§42.4
Thalamus plus hypothalamus
Thalamus§42.4
Primary sensory integration and relay station to the cerebral lobes
Hypothalamus§42.4
Regulates temperature, hunger, thirst, emotion; controls the pituitary
Telencephalon / cerebrum§42.4
Front forebrain devoted to association, correlation, and learning
Corpus callosum§42.4
Axon tract connecting the two cerebral hemispheres
Contralateral§42.4
On the opposite side of the body from the controlling hemisphere
Cerebral cortex§42.4
Few-mm gray-matter surface layer with >10 billion neurons
Gyrus / sulcus§42.4
A surface fold and a surface groove of the brain
Primary motor cortex§42.4
Frontal lobe strip controlling voluntary movement, mapped by body part
Primary somatosensory cortex§42.4
Parietal lobe strip receiving skin and muscle sensation, mapped by body part
Association cortex§42.4
Non-motor, non-sensory cortex for higher mental activity; ~95% in humans
Basal nuclei§42.4
Gray-matter islands deep in cerebral white matter that help control movement
Limbic system§42.4
Hippocampus, amygdala, and hypothalamus — emotion and memory
Hippocampus§42.4
Limbic structure for forming and consolidating memories
Reticular activating system (RAS)§42.4
Brainstem network controlling alertness, consciousness, and sleep–wake state
EEG§42.4
Recording of brain electrical activity; alpha, beta, theta, delta waves
Lateralization§42.4
Specialization of the two hemispheres for different functions
Broca’s area§42.4
Frontal lobe area generating the motor output of speech
Wernicke’s area§42.4
Parietal lobe area for language comprehension; damage gives word salad
Aphasia§42.4
Language disorder caused by damage to a language area
Prosopagnosia§42.4
Inability to recognize faces, from nondominant inferior temporal damage
Long-term potentiation (LTP)§42.4
Synapse strengthened by AMPA receptor insertion after strong NMDA activation
Long-term depression (LTD)§42.4
Synapse weakened by AMPA receptor removal via calcineurin
NMDA receptor§42.4
Glutamate receptor needing both ligand binding and depolarization (Mg²⁺ block removal)
Beta-amyloid / tau§42.4
Alzheimer suspects — external plaques and internal neurofibrillary tangles
Meninges§42.4
Protective membrane layers covering the brain and spinal cord
Reflex§42.4
Fast, involuntary muscle response processed at the spinal cord
Monosynaptic reflex arc§42.4
Reflex with one synapse, sensory straight to motor (knee-jerk)
Cutaneous reflex§42.4
Skin reflex routed through one or more interneurons
Dorsal root§42.5
Sensory branch of a spinal nerve entering the cord dorsally
Ventral root§42.5
Motor branch of a spinal nerve leaving the cord ventrally
Dorsal root ganglion§42.5
Cluster of sensory neuron cell bodies just outside the spinal cord
Preganglionic neuron§42.5
First autonomic motor neuron; CNS cell body, always releases ACh
Postganglionic neuron§42.5
Second autonomic motor neuron; ganglion cell body to the effector organ
Nicotinic receptor§42.5
ACh receptor on skeletal muscle that opens directly as a Na⁺ channel
Muscarinic receptor§42.5
G-protein-linked ACh receptor; opens heart K⁺ channels and slows the beat
G protein§42.5
Membrane protein activated by a receptor that then activates an effector protein
Sympathetic chain§42.5
Paired ganglion chains beside the cord holding sympathetic postganglionic cell bodies
Adrenal medulla§42.5
Inner adrenal region secreting epinephrine when sympathetic fibers fire
Cranial nerve§42.5
One of 12 paired nerves arising from the underside of the brain, I–XII
Vagus nerve (X)§42.5
Cranial nerve reaching chest and abdominal organs; central to homeostasis
Mixed nerve§42.5
Nerve carrying both sensory and motor fibers